Figure 1. Burrowing talitrids create a well-defined bioturbation zone on gently sloping sandy beaches in southern California. (a) Location of study site. (b) The burrowing zone (black dashed line) on Isla Vista Beach on a typical summer’s day. (c) Photograph of Megalorchestia spp. provided by Nicholas Schooler. (d) Rectangular metal frames used to estimate the mounding rate of burrowing talitrids. (e) Diagram showing the spatial arrangement of the 21 sampling plots (not to scale).